ƒ_ƒCƒZƒ“ƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Daisenialoe)
@ @ ƒ_ƒCƒZƒ“ƒiƒKƒSƒ~ƒ€ƒV Pterostichus fujimurai ’†‘`‹ß‹E–k•” (”ŒãË‘åŽR)
@ ƒiƒ“ƒLƒiƒKƒSƒ~ƒ€ƒV Pterostichus shotaroi ‹IˆÉ”¼“‡ (“ú’uì’¬‰ÍŒ´’J)
ƒqƒ‰ƒ^ƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Sphodroferonia)
@ ƒVƒRƒNƒqƒ‰ƒ^ƒiƒKƒSƒ~ƒ€ƒV Pterostichus miyamai Î’ÈŽRŒn (‘éƒm‘ƒŽR)
ƒqƒ‡ƒEƒSƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Pterostichus)
ƒjƒbƒRƒEƒIƒIƒYƒiƒKƒSƒ~ŽíŒQ
@ ƒjƒbƒRƒEƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus macrogenys “Œ–k`ŠÖ“Œ`’†•”
@ ƒGƒ`ƒSƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus asahinus ’©“úŠx
@ ƒ„ƒcƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus koheii ”ªƒ–Šxi–kŠÖ“ŒE‘å•ìŽFj
@ ƒAƒLƒ^ƒRƒ}ƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus tanakai H“c‹î‚ªŠx
@ ƒVƒRƒNƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus yoshidai Œ•ŽR
@ ƒqƒƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus toyodai ɪŒ§ˆ¢”{“»
@ ƒEƒGƒ_ƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus uedaorum ˆã‰¤ŽRA”’ŽR
@ ƒGƒiƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus todai ’·–쌧_â“»
ƒcƒVƒ}ƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒcƒVƒ}ƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus opacipennis ‘Δn
@ ƒqƒRƒTƒ“ƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus macrocephalus ‰p•FŽRi”wUE‘½—ÇE‹ãBŽR’nj
@ ƒSƒgƒEƒIƒIƒYƒiƒKƒSƒ~ƒ€ƒV Pterostichus amanoi ŒÜ“‡•Ÿ]“‡
@ ƒCƒLƒcƒLƒiƒKƒSƒ~ƒ€ƒV Pterostichus sakagamii ’·èŒ§¶ŒŽ“‡
ƒVƒCƒoƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒVƒCƒoƒiƒKƒSƒ~ƒ€ƒV Pterostichus shiibanus ’Å—t‘ºi‹ãBŽR’nj
@ @ ƒCƒVƒYƒ`ƒiƒKƒSƒ~ƒ€ƒV Pterostichus ishizuchiensis Î’ÈŽRŒn
ƒ€ƒiƒrƒƒiƒKƒSƒ~ƒ€ƒVŽíŒQ

@

 ƒ€ƒiƒrƒƒiƒKƒSƒ~ƒ€ƒV Pterostichus abaciformis ’†•”`‹ß‹E`’†‘ (“‡X’Jj
@ ƒ„ƒcƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus mucronatus ”ªƒ–Šxi‘½–€E’•ƒE‘å•ìŽFj
@ ƒ}ƒXƒ‚ƒgƒiƒKƒSƒ~ƒ€ƒV Pterostichus masumotoi “ìƒAƒ‹ƒvƒXi•xŽmŽRj
@ ƒgƒ_ƒCƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus kosakai ‰Ìh
ƒqƒ‡ƒEƒSƒiƒKƒSƒ~ƒ€ƒVŽíŒQ

@

 ƒqƒ‡ƒEƒSƒiƒKƒSƒ~ƒ€ƒV Pterostichus sphodriformis ‹ß‹E`’†‘
@ ƒWƒ…ƒ“ƒTƒCƒiƒKƒSƒ~ƒ€ƒV Pterostichus pachinus ŠÖ“Œ`’†•”iäñØÀj
@ ƒXƒ‹ƒKƒiƒKƒSƒ~ƒ€ƒV Pterostichus suruganus ɪŒ§•xŽm
@ ƒcƒ‹ƒKƒiƒKƒSƒ~ƒ€ƒV Pterostichus tsurugaensis •ŸˆäŒ§“Ö‰ê
@ ƒGƒTƒLƒiƒKƒSƒ~ƒ€ƒV Pterostichus esakii •CŒ©
@ ƒqƒTƒ}ƒcƒiƒKƒSƒ~ƒ€ƒV Pterostichus hisamatsui Î’ÈŽRŒn
@ ƒLƒCƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus pseudopachinus ‹ß‹E`’†‘i‘å‘䃖Œ´j
@ ƒIƒLƒiƒKƒSƒ~ƒ€ƒV Pterostichus okiensis ‰BŠò
@ ƒSƒgƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus gotoensis ’·èŒ§ŒÜ“‡—ñ“‡
@ ƒqƒRƒTƒ“ƒiƒKƒSƒ~ƒ€ƒV Pterostichus hikosanus ‹ãBŽR’nAŽl‘¼•”
@ ƒIƒJƒ„ƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus nasui ‰ªŽR
ƒLƒ…ƒEƒVƒ…ƒEƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒLƒ…ƒEƒVƒ…ƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus kyusyuensis ‹ãBŽR’nAŽl‘
@ ƒˆƒVƒJƒƒiƒKƒSƒ~ƒ€ƒV Pterostichus yoshikawai ‹IˆÉ”¼“‡i‘å‘䃖Œ´j
@ ƒ}ƒVƒ_ƒiƒKƒSƒ~ƒ€ƒV Pterostichus masidai ’†‘iŠ¥ŽRj
@ ƒCƒgƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus itoi Î’ÈŽRŒn
@ ƒjƒZƒLƒ…ƒEƒVƒ…ƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus arcuaticarinatus ‹ãB
@ ƒCƒ}ƒTƒJƒiƒKƒSƒ~ƒ€ƒV Pterostichus imasakai ‘½—ÇŠxi‰_åŠxj
@ ƒ^ƒ‰ƒ_ƒPƒiƒKƒSƒ~ƒ€ƒV Pterostichus taradakensis ‘½—ÇŠxi”wUŽRj
@ ƒ\ƒ{ƒiƒKƒSƒ~ƒ€ƒV Pterostichus fenestratus ‘c•êŽR
@ ƒ„ƒ}ƒWƒiƒKƒSƒ~ƒ€ƒV Pterostichus yamajii ‰ªŽRi’†˜a‘ºŽRæŽRj
ƒgƒTƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒgƒTƒiƒKƒSƒ~ƒ€ƒV Pterostichus tosanus Î’ÈŽRŒn (“y²’¬Š~ŽR)
ƒnƒ‰ƒgƒQƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Epinialoe)
@ ƒnƒ‰ƒgƒQƒiƒKƒSƒ~ƒ€ƒV Pterostichus spiculifer “Œ–k`ŠÖ“Œ`’†•”
@ ƒ„ƒcƒnƒ‰ƒgƒQƒiƒKƒSƒ~ƒ€ƒV Pterostichus yatsuensis ”ªƒ–Šx
@ ƒVƒ“ƒVƒ…ƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus cristatoides –kƒAƒ‹ƒvƒXi”’ŽRj
@ ƒ„ƒmƒiƒKƒSƒ~ƒ€ƒV Pterostichus janoi –kE’†‰›ƒAƒ‹ƒvƒX`”’ŽR
@ ƒ_ƒ“ƒhƒiƒKƒSƒ~ƒ€ƒV Pterostichus dandonis ’iŒË’J
@ ƒlƒIƒ_ƒjƒiƒKƒSƒ~ƒ€ƒV Pterostichus ohbayashii Šò•ŒŒ§ª”ö’J
@ ƒ_ƒCƒqƒUƒ“ƒiƒKƒSƒ~ƒ€ƒV Pterostichus daihizanus ‹ž“sEŽ ‰êE•Ÿˆä
@ ƒnƒNƒTƒ“ƒiƒKƒSƒ~ƒ€ƒV Pterostichus hakusanus ”’ŽR
ƒEƒGƒmƒIƒIƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Paralianoe)
ƒEƒGƒmƒIƒIƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒEƒGƒmƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus uenoi ŠÖ“Œ`’†•”`‹ß‹E (ã‚’n)
ƒ_ƒCƒZƒ“ƒ~ƒYƒMƒƒiƒKƒSƒ~ƒ€ƒV Pterostichus daisenicus ‹ß‹E`’†‘ (‘åŽRj
@ ƒ`ƒ…ƒEƒWƒ‡ƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus chujoi Œ•ŽR
@ ƒTƒgƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus satoi Î’ÈŽRŒn
@ ƒ^ƒJƒnƒVƒiƒKƒSƒ~ƒ€ƒV Pterostichus bisetosus ‹IˆÉ”¼“‡ (‘å‘䃖Œ´)
ƒŠƒ…ƒEƒWƒ“ƒiƒKƒSƒ~ƒ€ƒV Pterostichus tanakaorum ˜a‰ÌŽRŒ§—´_‘ºEŒì–€’dŽR
@ ƒVƒ‰ƒnƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus kiiensis ˜a‰ÌŽRŒ§”’•l’¬
ƒIƒIƒ_ƒCƒIƒIƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒIƒIƒ_ƒCƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus biexcisus ‹IˆÉ”¼“‡i‘å‘䃖Œ´j
@ ƒ_ƒCƒZƒ“ƒIƒIƒiƒKƒSƒ~ƒ€ƒV P. biexcisus subparalleles ’†‘AŽ]ŠòŽR’n
@ ƒJƒƒmƒiƒKƒSƒ~ƒ€ƒV Pterostichus kawanoi Œ•ŽR
ƒcƒ‹ƒMƒ„ƒ}ƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒcƒ‹ƒMƒ„ƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus tsurugiyamanus Œ•ŽR
@ ƒVƒƒ€ƒlƒIƒIƒiƒKƒSƒ~ƒ€ƒV Pterostichus omogoensis Î’ÈŽRŒn
@ ƒeƒ“ƒOƒiƒKƒSƒ~ƒ€ƒV Pterostichus carsticus “V‹ç‚Œ´
ƒ~ƒYƒMƒƒiƒKƒSƒ~ƒ€ƒVˆŸ‘® (Nialoe)
ƒ~ƒYƒMƒƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒ~ƒYƒMƒƒiƒKƒSƒ~ƒ€ƒV Pterostichus asymmetricus ’†•”`ŠÖ“Œ`“Œ–ki“úŒõj
@ ƒgƒPƒWƒiƒKƒSƒ~ƒ€ƒV Pterostichus tokejii ‰œ‘½–€A‰œ’•ƒiŽO•ôŽRj
@ ƒˆƒRƒnƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus yokohamae ‰¡•lŽs’ߌ©ì
@ ƒAƒuƒNƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus nishiyamai ˆ¢•ŒGŽR’ni”ªaŽRj
@ ƒTƒCƒgƒiƒKƒSƒ~ƒ€ƒV Pterostichus masahiroi •ŸˆäŒ§Š¥ŽR
@ ƒ€ƒTƒVƒiƒKƒSƒ~ƒ€ƒV Pterostichus musasiensis é‹Ê`“Œ‹ž`ŽR—œ‚Ì’áŽR’n
@ ƒCƒXƒ~ƒiƒKƒSƒ~ƒ€ƒV Pterostichus isumiensis ç—tŒ§ˆÎ‹÷’¬
ƒIƒNƒ^ƒ}ƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒIƒNƒ^ƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus okutamae ‰œ‘½–€iŽO“ªŽRj
@ ƒ^ƒ“ƒUƒƒiƒKƒSƒ~ƒ€ƒV Pterostichus watanabei ’O‘òŽR’ni•¨Œ©“»j
@ ƒ~ƒgƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus mitoyamanus ‰œ‘½–€A’O‘òiŽO“ªŽRj
ƒJƒ^ƒVƒiƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒJƒ^ƒVƒiƒiƒKƒSƒ~ƒ€ƒV Pterostichus katashinensis ŠÖ“ŒŽR’niŒQ”nŒ§•Ð•i‘ºj
@ ƒiƒJƒlƒiƒKƒSƒ~ƒ€ƒV Pterostichus nakanei –kE’†‰›ƒAƒ‹ƒvƒX
@ ƒŠƒ…ƒEƒgƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus tahirai ɪŒ§—³“ªŽRE‹¼”ž—±ŽR
@ ƒIƒKƒiƒKƒSƒ~ƒ€ƒV Pterostichus ogaensis “È–ØŒ§’jŽ­Šx
ƒ~ƒ„ƒ}ƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒ~ƒ„ƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus rhanis ŠÖ“ŒŽR’n`”ªƒ–Šx
@ ƒtƒNƒCƒiƒKƒSƒ~ƒ€ƒV Pterostichus cordatissimus •Ÿˆä`‹ß‹E`’†‘“Œ•”
@ ƒnƒlƒAƒJƒiƒKƒSƒ~ƒ€ƒV Pterostichus brunneipennis –kƒAƒ‹ƒvƒXA“ìƒAƒ‹ƒvƒX
@ ƒzƒYƒ~ƒiƒKƒSƒ~ƒ€ƒV Pterostichus hozumii ˆ¤’mŒ§’iŒËŽR
@ ƒtƒWƒiƒKƒSƒ~ƒ€ƒV Pterostichus fujisanus “ìƒAƒ‹ƒvƒXi•xŽmŽRj
@ ƒIƒIƒ_ƒCƒiƒKƒSƒ~ƒ€ƒV Pterostichus ohdaisanus ‹IˆÉ”¼“‡i‘å‘䃖Œ´j
@ ƒRƒ“ƒSƒEƒiƒKƒSƒ~ƒ€ƒV Pterostichus kongosanus ‹à„ŽR
@ ƒLƒ\ƒiƒKƒSƒ~ƒ€ƒV Pterostichus uchiyamai ’†‰›ƒAƒ‹ƒvƒXi‹{“c‚Œ´j
@ ƒAƒJƒVƒiƒiƒKƒSƒ~ƒ€ƒV Pterostichus ishizukai ’·–쌧–¾‰È’¬
ƒ^ƒiƒJƒiƒKƒSƒ~ƒ€ƒVŽíŒQ
@ ƒ^ƒiƒJƒiƒKƒSƒ~ƒ€ƒV Pterostichus latistylis “Œ–k`ŠÖ“Œ`’†•”`’†‘
@ ƒVƒoƒ^ƒiƒKƒSƒ~ƒ€ƒV Pterostichus shibatai ‹IˆÉ”¼“‡iŒì–€’dŽRj
@ ƒIƒIƒNƒ‰ƒiƒKƒSƒ~ƒ€ƒV Pterostichus ohkurai –kƒAƒ‹ƒvƒXi‰_ƒm•½j
@ ƒLƒ\ƒRƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus napaea –Ø‘]‹îƒ–Šx
@ ƒZƒgƒiƒKƒSƒ~ƒ€ƒV Pterostichus taoi ˆ¤’mŒ§£ŒËŽs
ƒtƒ^ƒgƒQƒiƒKƒSƒ~ƒ€ƒVŒQiˆŸ‘®–¢’èj
@ ƒtƒ^ƒgƒQƒiƒKƒSƒ~ƒ€ƒV Pterostichus mirificus “Œ–k`“n“‡”¼“‡
ƒcƒVƒ}ƒiƒKƒSƒ~ƒ€ƒVŒQiˆŸ‘®–¢’èj
@ ƒcƒVƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus symmetricus ‘Δn
ƒ„ƒNƒVƒ}ƒiƒKƒSƒ~ƒ€ƒVŒQiˆŸ‘®–¢’èj
@ ƒ„ƒNƒVƒ}ƒiƒKƒSƒ~ƒ€ƒV Pterostichus yakushimanus ‰®‹v“‡
ƒAƒ}ƒ~ƒiƒKƒSƒ~ƒ€ƒVŒQiˆŸ‘®–¢’èj
@ ƒAƒ}ƒ~ƒiƒKƒSƒ~ƒ€ƒV Pterostichus plesiomorphs ‰‚”ü‘哇“’˜pŠx