Goeridae

A catalogue of Japanese Trichoptera
21. Family Goeridae Ulmer

Author: Takao NOZAKI
E-mail: takao.nozaki[AT]nifty.com
Last update: Mar. 1, 2008

Subfamily Goerinae Ulmer

Two genera, Goera and Silo, were recorded from Japan, but identification of Silo is very doubtful (Nozaki et al. 2000)

Genus Goera Stephens

Fifteen species belonging to this genus are recognized for Japanese fauna (Nozaki and Tanida 2006).

Goera akagiae Tanida and Nozaki 2006

Goera sp. GC: Akagi 1974, 18-19, larva, case, male, Amami-Oshima. Associated with the adult stage by Nozaki and Tanida (2006).

Tanida 1985, 196, 197, larva.
Nozaki et al. 2000, 198.
Nozaki 2005, 536, 537, larva.

Goera akagiae: Nozaki and Tanida 2006, 7-9, male, female, Shikoku (Kochi), Yaku-shima, Amami-Oshima.
Type locality. Amami-Oshima, Japan.
Distribution. Japan (Shikoku, Yaku-shima, Amami-Oshima).

Goera curvispina Martynov 1935

Goera curvispina Martynov 1935, 367-370, male.

Tsuda and Akagi 1955, 237, Honshu (Shiga).
Akagi 1975, 3-4, larva, case, Honshu (Yamaguchi), Kyushu (Fukuoka, Oita).
Gyotoku and Nozaki 1992, 138, Honshu (Fukuoka).
Fischer 1973, 1.
Tani 1976, 393, Tsushima (larva).
Tani 1977, 205, male.
Arefina 1997, 127-128, male.
Cho et al. 1999, 46, male.
Nozaki et al. 2000, 198.
Uenishi and Yoshitaka 2004, 17, Honshu (Okayama).
Nozaki 2005, 535-538, male, larva.
Nozaki and Tanida 2006, 10-12, male, female, Honshu (Hyogo, Shimane), Kyushu (Fukuoka), Russia.

Goera gyotokui Kobayashi 1957, 276-278, male, Fukuoka. Synonymized by Nozaki and Tanida (2006).

Fischer 1973, 2.
Nozaki et al. 2000, 198.

Goera gyotokuana [!]: Kobayashi 1959, 354, Fukuoka.
Type locality. South Primorye (Ussuri R.), Russia.
Distribution. Japan (Honshu (central to western), Kyushu, Tsushima), Korea, Russia (Khabarovsk, Primorye).

Goera dilatata Nozaki and Tanida 2006

Goera dilatata Nozaki and Tanida 2006, 25-27, male, female, Hokkaido (Oshima), Honshu (Iwate, Miyagi, Akita, Niigata).
Type locality. Honshu (Akita), Japan.
Distribution. Japan (Hokkaido (southern), Honshu (northern))

Goera japonica Banks 1906

Goera japonica Banks 1906, 108-109, male, Honshu (Shizuoka?).

Ota 1917, 489-494, parasitoid, Honshu (Kanagawa).
Matsumura 1931, 191, adult.
Matsumura 1933, 2, pl. 1, adult.
Tateishi 1934, 75-76, Kyushu (Kagoshima).
Tsuda 1941, 104, Honshu (Nagano).
Tsuda 1942a, 323-324, Honshu (Nagano, Shizuoka, Kyoto, Osaka, Hyogo), Kyushu (Fukuoka).
Tsuda 1942b, 263-265, Honshu (Kyoto).
Tsuda 1943, 163166-168, Honshu (Kyoto).
Tsuda and Kuwayama 1950, 428, adult.
Tsuda 1951, 66, larva, case.
Ishihara et al. 1953, 38, Shikoku (Ehime).
Tsuda and Akagi 1955, 235-236, larva, case.
Chihara 1956, 81-82, pupa.
Gyotoku 1957, 12-13, Kyushu (Fukuoka).
Miwa and Ohkawa 1958, 19, Honshu (Mie).
Kobayashi 1959, 353-354, Kyushu (Fukuoka).
Tsuda 1959, 150, larva, case.
Tsuda and Akagi 1962, 140, larva, case.
Kuwada 1965, 166, pl.. 83, adult, Shikoku (Ehime).
Fischer 1967, 7.
Kobayashi 1968, 6, Honshu (Niigata).
Kobayashi 1971, 35-36, male, female, Honshu (Kanagawa).
Foscher 1973, 2.
Tani 1976, 393, Tsushima.
Tani 1977, 194, 205, pl. 44, male, Honshu (Nara).
Tani 1978, 16-17, Honshu (Niigata).
Tanida 1980, 136, Honshu (Ishikawa).
Tanida and Takemon 1981, 206, Honshu (Kyoto).
Tani et al. 1982, 34-35, Oki (based on larva).
Tanida 1982a, 361, Honshu (Gumma).
Tanida 1982b, 18-20, Honshu (Ishikawa).
Kobayashi 1983, 32, Honshu (Kanagawa.)
Kobayashi 1984a, 20, male, female, Hokkaido (Shiribeshi, Hiyama, Oshima).
Kobayashi 1984b, 7-8, Sado.
Kobayashi 1984c, 28, Honshu (Kanagawa).
Kobayashi 1985a, 19, Tsushima.
Kobayashi 1985b, 68, Honshu (Kanagawa).
Tanida 1985, 196, 197, larva, case.
Togashi and Tanida, 1985, 69, Honshu (Ishikawa).
Ohtsuka and Tanida. 1986, 6, Kyushu (Kumamoto).
Kobayashi 1987, 23-24, 1987, Honshu (Shimane).
Yamanouchi 1987, 36, Hokkaido (Tokachi).
Nozaki 1987, 41-42, Honshu (Kanagawa).
Morita 1988, 5, Honshu (Mie).
Nozaki 1988a, 45-46, Honshu (Kanagawa).
Nozaki 1988b, 42, Honshu (Kanagawa).
Morita 1989, 40, Honshu (Mie).
Nozaki and Gyotoku 1990, 17, Kyushu (Fukuoka).
Kugo and Nozaki 1990, 63, Honshu (Hyogo).
Morita 1990, 31, Honshu (Mie).
Tanida 1990, 56, Honshu (Hyogo).
Aoyagi and Ishii 1991, 113-118, parasitoid.
Hatta and Nozaki, 1991, 204, Honshu (Gifu).
Kugo and Nozaki, 1991, 8, Honshu (Hyogo).
Tanida 1991, 35, Honshu (Nara).
Tanida et al. 1991, 100-101, adult, pupa, larva, case.
Gyotoku and Nozaki 1992, 137-138, Kyushu (Fukuoka).
Moriya 1992, 44, 63, adult, case, Honshu (Kanagawa).
Hori et al. 1993, 375, 389, Honshu (Aichi).
Kobayashi 1993, 401-402, parasitoid.
Kuhara et al. 1993, 18, Hokkaido (Ishikari).
Morita 1993, 6, Honshu (Mie).
Nishimoto and Nishimoto 1993, 87, Honshu (Aichi).
Tanida and Takemon 1993, 243, Honshu (Kyoto)
Kuhara and Ito 1994, 21, Hokkaido (Ishikari).
Kuranishi and Kuhara 1994, 1229, Hokkaido (Akan).
Moriya 1994, 79, Honshu (Kanagawa, Yamanashi).
Kobayashi 1995, 318-321, parasitoid.
Tanida et al. 1995, 3, Honshu (Nara).
Kuranishi and Ohtaka 1996, 15, Honshu (Aomori).
Kuhara 1997a, 61, Hokkaido (Shiribeshi).
Kuhara 1997b, 96, Hokkaido (Shiribeshi)
Nakajima et al. 1997, 14, Hokkaido (Oshima).
Ito et al. 1997, 26, 28, 30, 31, Hokkaido (Hiyama).
Morita 1997a, 41, Honshu (Mie).
Morita 1997b, 96, Honshu (Mie).
Moriya 1997, 68, Honshu (Kanagawa, Yamanashi).
Nozaki 1997a, 36, Honshu (Kanagawa).
Nozaki 1997b, 30, Honshu (Tokyo).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Kadowaki 1998, 8, Honshu (Shimane).
Kagaya et al. 1998, 29, 61, 199, Honshu (Tokyo, Yamanashi).
Kobayashi, 1998, 109-113, parasitoid.
Ito et al. 1998, 12, 13, 15, 16, Hokkaido (Shibetsu).
Morita 1998a, 14, Honshu (Mie).
Morita 1998b, 73, Honshu (Mie).
Ito et al. 1999, 25-27, Hokkaido (Akan).
Nakano and Tanida 1999, 277, Honshu (Nara).
Ohkawa 1999, 41-44, Hokkaido (Tokachi).
Imamori 2000, 234-235, adult, larva.
Ito et al. 2000a, 27-31, Hokkaido (Soya).
Ito et al. 2000b, 35, Hokkaido (Ishikari).
Kuhara et al. 2000, 255, Hokkaido (Iburi).
Maruyama and Takai 2000, 108, photo 2-25, larva, case, Shikoku (Kochi).
Nishimoto 2000, 26, Honshu (Aichi).
Nozaki et al. 2000, 198.
Uenishi and Ebisawa 2000, 170, Honshu (Shiga).
Kawase and Morita 2001, 9, Honshu (Ishikawa).
Kuhara 2001, 19, Hokkaido (Shiribeshi).
Morita 2001a, 221, Honshu (Aichi).
Morita 2001b, 164, Hosnhu (Mie).
Morita 2001c, 339, Honshu (Mie).
Nishimoto and Morita 2001, 77, Hosnhu (Aichi).
Kuranishi 2002, 571, larva, Honshu (Chiba).
Nakamura et al. 2002, 67, Honshu (Hiroshima).
Nozaki and Nakamura 2002, 173-174, Honshu (Hiroshima).
Karita 2003, 130-131, adult, larva, case.
Nagayasu et al. 2003, 110, Hokkaido (Ishikari).
Nakamura and Umemori 2003, 59-60, Honshu (Hiroshima).
Shinoda 2003, 145, Honshu (Yamanashi).
Nakamura et al. 2004, 154, Honshu (Hiroshima).
Kawase et al. 2004, 114, Honshu (Ibaraki)
Morita and Kawase 2004, 16, 18, Honshu (Ishikawa).
Nozaki 2004, 1236, Honshu (Kanagawa).
Taira and Kuranishi 2004, 74, Honshu (Chiba).
Yoshitaka 2004, 28, Honshu (Okayama).
Katsuma 2005a, 7, Honshu (Ibaraki).
Katsuma 2005b, 14, Honshu (Tochigi, Gumma).
Nagayasu et al. 2005, 56, Hokkaido (Ishikari).
Nakano 2005, 305-308, pupal aggregation, Hokkaido (Ishikari).
Nozaki 2005, 535-538, male, larva, case.
Nozaki 2005, 112, adult, larva, case.
Nozaki and Nakamura 2005, 224, Honshu (Shimane).
Kameyama and Nakamura 2006, 190, Honshu (Tottori, Hiroshima).
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Katsuma 2006, 41, Honshu (Ibaraki).
Kuhara et al. 2006, 46, Hokkaido (Ishikari).
Mey and Nozaki 2006, 25, Honshu (Nagano, Gifu, Shiga).
Morita 2006a 201, Honshu (Mie).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Nozaki and Tanida 2006, 3-6, male, female, Hokkaido (Nemuro, Abashiri, Soya, Kamikawa, Hiyama), Honshu (Miyagi, Akita, Fukushima, Tochigi, Gumma, Tokyo, Kanagawa, Niigata, Nagano, Gifu, Shizuoka, Aichi, Mie, Hyogo, Nara, Hiroshima, Shikoku (Tokushima, Ehime, Kochi), Sado, Tsushima, Yaku-shima.
Torii and Hattori 2006, 39, Honshu (Shizuoka).
Gall et al. 2007, 108-111, male, female, wings of female, Hokkaido (Iburi), Honshu (Kanagawa), Sakhalin, Kuriles, Cheju (South Korea).
Ito and Kosugi 2007, 55, Hokkaido (Kushiro).
Katsuma 2007, 18, Honshu (Ibaraki).
Kimura et al. 2007, 167, Honshu (Nagano).
Kuhara et al. 2007, ap. 72, Hokkaido (Ishikari).
Morita 2007a, 15, Honshu (Mie).
Morita 2007b, 97, Honshu (Mie).
Nakano et al. 2007, 41-42, Hokkaido (Nemuro).
Nakano et al. 2007, 103-110, effects by case aggregation, Hokkaido (Ishikari).
Nozaki and Nakamura 2007, 98, Honshu (Hiroshima).
Nozaki and Tanida 2007, 251, Honshu (Shizuoka).
Shinoda 2007, 97, Honshu (Yamanashi).

Lithax karafutonis Matsumura 1911, 16, female (as male in the original description), Sakhalin. Synonymized by Gall et al. 2007).
Goera pilosa Fabricius: Iwata 1927a, 255-256, larva. Misidentification mentioned by Tsuda and Akagi (1955).

Iwata 1927b, 214, larva.
Fischer 1967, 15-16.

Goera nipponensis Navas 1933, 107-108, male, female, Honshu (Yamanashi). Synonymized by Nozaki and Tanida (2006).

Fischer 1967, 8.
Nozaki et al. 2000, 198.

Type locality. Honshu (Kawana: Shizuoka?), Japan.
Distribution. Japan (Hokkaido, Honshu, Shikoku , Kyushu, Sado, Oki (based on larva), Tsushima, Yaku-shima), Russia (Sakhalin, Kuriles), Korea (Cheju).
Remarks. Gall et al. (2007) resolved the taxonomic status of G. japonica and similar species from the eastern Palaearctic Region.

Goera kawamotonis Kobayashi 1987

Goera sp. GA, Tsuda and Akagi 1956, 39-40, larva, case, male, Honshu (Aichi). Associated with the adult stage by Nishimoto and Morita (2001).

Tsuda and Akagi 1962, 140, larva.
Fischer 1973, 5.
Tanida 1985, 196, 199, larva.
Nozaki et al. 2000, 198.

Goera kawamotonis Kobayashi 1987, 24, 34, male, Honshu (Shimane).

Gyotoku and Nozaki 1992, 138, Kyushu (Fukuoka).
Arefina 1997, 126, 128, male.
Nozaki et al. 2000, 198.
Nishimoto and Morita 2001, 77, Honshu (Aichi).
Morita 2002, 37, Honshu (Mie).
Nozaki and Nakamura 2004, 224, Honshu (Shimane).
Yoshitaka 2004, 28, Honshu (Okayama).
Uenishi and Yoshitaka 2004, 17-18, Honshu (Okayama).
Nozaki 2005, 535-538, male, larva, case.
Kameyama et al. 2006, 94, Honshu (Hiroshima).
Nakamura and Fukui 2006, 244, Honshu (Hiroshima).
Nozaki and Tanida 2006, 11-13, male, female, Honshu (Aichi, Hyogo, Hiroshima, Shimane, Yamaguchi), Shikoku, Kyushu (Fukuoka).
Nozaki and Nakamura 2007, 98, Honshu (Hiroshima).

Type locality. Honshu (Shimane), Japan.
Distribution. Japan (Honshu (central to western), Kyushu), Russia (Khabarovsk, Primorye).
Remarks. Nozaki and Tanida (2006) added Shikoku as a distribution area, but no data was provided.

Goera kyotonis Tsuda 1942

Goera kyotonis Tsuda 1942a, 324-325, male, Honshu (Kyoto).

Tsuda and Akagi 1955, 236-237, larva, case, Honshu (Kyoto).
Kobayashi 1957, 278, pl. 44, male.
Tsuda and Akagi 1962, 140, larva, case.
Tsuda 1959, 150, larva, case.
Fischer 1967, 7.
Fischer 1973, 3.
Tani 1982, 34-35, Oki.
Morita 1996, 289, Honshu (Mie).
Tani and Nakamura 1997, 64, Honshu (Hiroshima).
Nozaki et al. 2000, 198.
Nozaki 2005, 535-538, male, larva, case.
Nozaki and Tanida 2006, 13-15, male, female, Honshu (Osaka, Hyogo, Wakayama, Tottori), Kyushu (Fukuoka).
Morita 2006b 47, Honshu (Mie).

Type locality. Honshu (Kyoto), Japan.
Distribution. Japan (Honshu (central to western), Kyushu, Oki (based on larva)).

Goera lepidoptera Schmid 1965

Goera lepidoptera Schmid 1965, 35, male, Honshu (Nagano).

Schmid 1991, 325, male maxillary palpus.
Kagaya et al. 1998, 29, 61, 200, Honshu (Tokyo).
Nozaki et al. 2000, 198.
Nozaki 2004, 1236, Honshu (Kanagawa)
Nozaki 2005, 538, male.
Katsuma 2006, 36, 41, Honshu (Ibaraki).
Nozaki and Tanida 2006, 15-16, male, female, Honshu (Miyagi, Yamagata, Kanagawa).

Goera kyotonis Tsuda: Moriya 1992, 44, Honshu (Kanagawa). Misidentification mentioned by Nozaki (2004).
Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu central to northern))
Remarks. Larvae of this species are indistinguishable from those of G. kyotonis (Nozaki and Tanida 2006).

Goera nigrosoma Nozaki and Tanida 2006

Goera sp. GB: Tsuda and Akagi 1962, 140, larva.

Tanida 1985, 196, 199, larva.
Tanida and Takemon 1993, 243, Honshu (Kyoto).
Kagaya et al. 1998, 29, 61, 202, Honshu (Tokyo).
Nozaki 2005, 536, 537, larva.
Nozaki et al. 2000, 198.

Goera sp.: Morita 1995, 49, male, Honshu (Mie). Reidentified as this species by Morita (2007).
Goera nigrosoma Nozaki and Tanida 2006, 20-22, male, female, Honshu (Kanagawa, Yamanashi, Nagano, Shizuoka, Kyoto, Osaka, Wakayama, Okayama), Shikoku (Kagawa, Ehime, Kochi).

Torii and Hattori 2006, 39, Honshu (Hokkaido).
Morita 2007, 15, Honshu (Mie).

Type locality. Honshu (Osaka), Japan.
Distribution. Japan (Honshu (central to western), Shikoku).

Goera ogasawaraensis Kuranishi 2005

Goera sp.: Tomokuni and Sato 1978, 117-118, larva, case, Ogasawara (Chichi-jima). Associated with the adult stage by Satake et al. (2005).

Nozaki et al. 2000, 198.

Goera ogasawaraensis: Satake et al. 2005, 376, 378-379, male, female, Ogasawara (Chichi-jima).

Nozaki and Tanida 2006, 5-6, male, female, Ogasawara (Chichi-jima).
Satake and Kuranishi 2007, 283-284, Ogasawara (Chichi-jima, Ani-jima).

Type locality. Ogasawara (Chichi-jima), Japan.
Distribution. Japan (Ogasawara (Chichi-jima, Ani-jima)).

Goera shikokuensis Nozaki and Tanida 2006

Goera shikokuensis Nozaki and Tanida 2006, 15-17, male, female, Shikoku (Ehime, Kochi).
Type locality. Shikoku (Ehime), Japan.
Distribution. Japan (Shikoku).

Goera spicata Schmid 1965

Goera spicata Schmid 1965, 35, male, Honshu (Nagano).

Schmid 1991, 325, male maxillary palpus.
Moriya 1994, 79, Honshu (Kanagawa, Yamanashi).
Moriya 1997, 68, Honshu (Kanagawa, Yamanashi).
Nozaki 1997, 36, Honshu (Kanagawa).
Kagaya et al. 1998, 29, 61, 201, Honshu (Tokyo, Yamanashi).
Nozaki et al. 2000, 198.
Nozaki 2004, 1236, Honshu (Kanagawa).
Nozaki 2005, 538, male.
Mey and Nozaki 2006, 25, Honshu (Nagano).
Nozaki and Tanida 2006, 24-26, male, female.

Type locality. Honshu (Nagano), Japan.
Distribution. Japan (Honshu (central)).
Remarks. Larval stage of this species was associated with adult and pupal stages by Kagaya et al. (1998), but the morphology has not yet been described.

Goera tajimaensis Tanida and Nozaki 2006

Goera tajimaensis: Nozaki and Tanida 2006, 22-23, male, female, Honshu (Hyogo).
Type locality. Hnshu (Hyogo), Japan.
Distribution. Japan (Honshu (central)). Known only from the type locality.

Goera tenuis Ulmer 1927

Goera tenuis Ulmer 1927, 181, male

Nozaki and Tanida 2006, 17-19, male, female, Ishigaki-jima, Iriomote-jima.

Type locality. Taiwan.
Distribution. Japan (Ishigaki-jima, Iriomote-jima), Taiwan.

Goera tungusensis Martynov 1909

Goera tungusensis Martynov, 1909, 246-250, male, female

Nozaki and Tanida 2006, 18-20, male, female, Hokkaido (Soya, Rumoi, Sorachi, Shiribeshi).

Goera sp. 1: Ito et al. 2000, 23, 27-29, Hokkaido (Soya). Reidentified as this species by Nozaki and Tanida (2006).
Type locality. Northern Siberia, Russia.
Distribution. Japan (Hokkaido), Russia (Siberia, Far East), Canada (Quebec), USA (Alaska).

Goera uchina Tanida and Nozaki 2006

Goera sp. GD: Tanida 1997, 449.

Tanida 2003, 379, male, larva.

Goera uchina: Nozaki and Tanida 2006, 8-10, male, female, Okinawa, Ishigaki-jima.
Type locality. Okinawa, Japan.
Distribution. Japan (Okinawa, Ishigaki-jima).
Remarks. Larvae of this species are indistinguishable from those of G. akagiae (Nozaki and Tanida 2006).

Genus Silo Curtis

A single species, Silo nigricellus Matsumura, is recorded from Hokkaido, but the generic identification is very doubtful (Nozaki et al. 2000).

Silo? nigricellus Matsumura 1907, Hokkaido

Silo nigricellus Matsumura 1907, 191, male.

Nozaki et al. 2000, 198.

Type locality. Hokkaido, Japan.
Distribution. Japan (Hokkaido).
Remarks. Only the color, size, and morphology of the antenna were provided in the original description by Matsumura (1907), but they are insufficient even for generic identification (Nozaki et al. 2000). The holotype of this species was not found in the collection of Matsumura deposited in Systematic Entomology, Faculty of Agriculture, Hokkaido University (Nozaki et al. 2000).

Subfamily Larcasinae Navas

Only the genus Larcasia belongs to this subfamily.

Genus Larcasia Navas

Two species of this genus were recorded from Japan (Nishimoto et al 1999).

Larcasia akagiae Nishimoto and Tanida 1999

Goerinae sp.: Akagi 1959, 42, larva, male, Shikoku (Ehime). Associated with the adult stage by Nishimoto et al. (1999)

Tsuda and Akagi 1962, 140, larva.

Imania sp. IA: Tanida 1985, 195, comb. Misidentification mentioned by Nishimoto et al. (1999).
Larcasia akagiae: Nishimoto et al. 1999, 426-435, male, female, pupa, larva, egg mass, case, habitat, adult behavior, life history, Honshu (Aichi, Nara).

Nishimoto 2000, 26-28, larva, adult, egg mass, habitat, adult behavior, Honshu (Aichi).
Nozaki et al. 2000, 199.
Nozaki 2005, 536-538, male, larva, case.

Type locality. Honshu (Aichi).
Distribution. Japan (Honshu (central), Shikoku).

Larcasia minor Nishimoto 1999

Larcasia minor: Nishimoto et al. 1999, 435-438, male, female, larva, egg mass, case, habitat, adult behavior, life cycle, Honshu (Tochigi).

Nozaki 2005, 536-538, male, larva.
Nozaki et al. 2000, 199.

Type locality, Honshu (Tochigi).
Distribution. Japan (Honshu (central)).

References

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